Symfony version(s) affected: 4.3.3

Description
I am trying to force a parameter being set in a route using!. It is working with the following annotation:

@Route("/titles/{!page}", name="list", methods={"GET"}, defaults={"page": 1})

When I generate the route without specifying a parameter ({{ path('list') }}) the following path is generated: /titles/1.

Unfortunately when combining !with other short syntax elements it does not work as expected:

• Specifying a default value using ?1:

  @Route("/titles/{!page?1}", name="list", methods={"GET"})
  

  produces: /titles/%7B!page%3F1%7D

• Specifying default as part of method signature:

  /**
   * @Route("/titles/{!page}", name="list", methods={"GET"})
   */
  public function list(int $page = 1) { ... }
  

  throws Twig\Error\RuntimeError:

  Some mandatory parameters are missing ("page") to generate a URL for route "list".

• Specifying requirements for page using short syntax <\d+>:

  @Route("/titles/{!page<\d+>}", name="list", methods={"GET"}, defaults={"page": 1})
  

  produces: /titles/%7B!page%3C%5Cd+%3E%7D

How to reproduce

1. Install annotations

   composer require annotations
   

2. Create a template with a link to the route using the force parameter: {{ path('...') }}
3. Create a controller action rendering the template with the route link
4. Create a controller action with one of the above route definitions containing !
5. Go to the initially defined route (template with link)